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Fulfill departmental requirements

For TA opportunities – serve one semester as a teaching assistant in Biophysics modules

Course Requirements

(term, # of credit hours in parentheses)

Required Track Cores

In addition to the departmental core courses (BIOC 704/710), complete the biophysics core courses listed below:

  • BIOC 649* Essentials of Macromolecular Science: Review basic principles of chemical & computational science, led by faculty/students (Fall, 1.5) *This course is only required for people who enter the program before 2018.
  • BIOC 650 Basic Principles: From basic models to collections of macromolecules (Fall, 1)
  • BIOC 651 Macromolecular Equilibria: Conformation change and binding (Fall, 1)
  • BIOC 652 Macromolecular Structure & Dynamics (Fall, 1)
  • BIOC 662 Macromolecular Interactions or BIOC 670 Structural Bioinformatics (Spring, 1) *This course is only required for students who matriculated prior to 2023.

Additional required course:

  • BCB888/BIOC 888 – Responsible conduct of research (1 credit): generally, fourth year in spring

Choose three from these modules:
Biophysics Program Modules:

  • BIOC 660 Imaging (Spring, 1)
  • BIOC 662 Macromolecular Interactions (Spring, 1)
  • BIOC 663A and 663B Macromolecular NMR and Theory (Spring, 1 ea)
  • BIOC 664 Macromolecular Spectroscopy: Principles and Applications (Spring, 1)
  • BIOC 665 Advanced NMR (Spring, 1)
  • BIOC 666 X-ray Crystallography of Macromolecules (Spring, 1)
  • BIOC 667 Macromolecular Crystallographic Methods (Spring, 1)
  • BIOC 669 Advanced Macromolecular Crystallographic Methods (Spring, 1)
  • BIOC 670 Structural Bioinformatics (Spring, 1)
  • BIOC 673 Proteomics, Protein Identification & Characterization by Mass Spec. (Spring, 1)
  • BIOC 674 Ion Channels Transporters (Spring, 1)
  • BIOC 675 Fundamentals of Cryo-Electron Microscopy (Spring, 1)
  • BCB 715: Mathematical and Computational Approaches to Modeling Signaling Pathways and Regulatory Networks (Fall, 1)
  • BMME 490/890: Super Resolution – imaging beyond limits (Fall, 1)
  • PHCO 750 Proteomics Methods and Applications (Spring, 1)


Complete a minimum of three credit hours from the options listed below:

  • Choose from any graduate-level course in a discipline supportive of biomedical research training, excluding the Biophysics track and Departmental core courses, and including
  • Molecular & Cell Biology
    BIOC 631 Advanced Molecular Biology I (Spring, 4)
    BIOC 706 Biochemistry of Human Disease (Fall, 3)
  • OR choose from these modules:
    BIOC 740-745 Contemporary Topics in Cell Signaling (Spring, 1) *740-742 offered in odd years and 743-745 in even years

*For students matriculating before 2023, a minimum of one elective credit hour is required


The course requirements for a certificate in Biophysics offered through the Molecular and Cellular Biophysics training program can be found on the training program website.

More information on the Graduate School’s Ph.D. Degree Requirements and Admission to Candidacy can be found in the Graduate School Handbook.